His death has pushed the northern white rhino to the very brink of extinction. Only two females of the subspecies now remain on the planet.
Sudan -- the last known male northern white rhino and probably the most popular male of his species -- died on March 19, 2018.
He is survived only by two females of the subspecies, putting a huge question mark on the future of the northern white rhino and the ability of humans to protect the species at large from our own kind.
As conservationists and all those who care about the environment mourn the loss of this magnificent animal, we look back at the life of Sudan.
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Sudan walks at the zoo in Dvůr Králové in the Czech Republic in 2009.
White rhinos are the largest land mammals after elephants and typically live in herds of up to 14 animals. The numbers of the northern subspecies have plummeted due mainly to poachers.
Sudan, who was born in 1973 in Sudan, will be remembered for his unusually memorable life.
Paying tribute to him, the Ol Pejeta Conservancy said, 'In the 1970s, he escaped extinction of his kind in the wild when he was moved to Dvůr Králové Zoo.'
Photograph: Petr Josek/Reuters.
Rhino keeper Berry White with Sudan at the zoo in Dvůr Králové.
Sudan and three other northern white rhinos were moved to the Ol Pejeta reservation in Kenya in 2009 as part of the 'Last Chance to Survive' breeding programme.
Photograph: Petr Josek/Reuters.
Sudan at the Ol Pejeta Conservancy in January 2010. He was 36 then.
He had already sired two calves by then.
Photograph: Noor Khamis/Reuters.
A warden massages Sudan at the Ol Pejeta Conservancy.
Beyond the breeding programme, conservationists also hoped Sudan would raise awareness of the plight of endangered animals and the need for community involvement in conservation.
Photograph: Thomas Mukoya/Reuters.
Wardens assist Sudan as he grazes.
The breeding programme with Sudan remained unsuccessful.
Photograph: Thomas Mukoya/Reuters.
A warden guards Sudan in May 2017.
He was guarded 24 hours a day to protect him from poaching. The protection detail also included horn-embedded transmitters, watchtowers, fences, drones, and guard dogs.
Photograph: Baz Ratner/Reuters.
Members of the Maasai Cricket Warriors with Sudan after playing against the British Army Training Unit cricket team during a charity tournament called the 'Last Male Standing' at the Ol Pejeta Conservancy.
The cricket tournament was hosted with hopes to raise awareness of the plight of endangered animals.
Photograph: Thomas Mukoya/Reuters.
Conservation efforts also involved putting Sudan on Tinder last year as a way to raise awareness and funds.
Since late 2017, Sudan was being treated for age-related complications that led to degenerative changes in muscles and bones combined with extensive skin wounds.
Ol Pejeta said in a statement, 'His condition worsened significantly in the last 24 hours; he was unable to stand up and was suffering a great deal. The veterinary team from the Dvůr Králové Zoo, Ol Pejeta and Kenya Wildlife Service made the decision to euthanize him.'
Photograph: Baz Ratner/Reuters.
A newly unveiled sculpture The Last Three -- a life-size Rhino sculpture created by Australian artists and conservationists Gillie and Marc, depicting Sudan, (bottom), his daughter Najiin, (middle) and granddaughter Fatu (top) -- in New York City, March 15, days before Sudan died.
Ol Pejeta said in a statement, 'Throughout his existence, he significantly contributed to survival of his species as he sired two females. Additionally, his genetic material was collected and provides a hope for future attempts at reproduction of northern white rhinos through advanced cellular technologies.'
They added, 'The only hope for the preservation of this subspecies now lies in developing in vitro fertilisation (IVF) techniques using eggs from the two remaining females, stored northern white rhino semen from males and surrogate southern white rhino females.'
Photograph: Mike Segar/Reuters.
Richard Vigne, Ol Pejeta's CEO said in a statement, 'He was a great ambassador for his species and will be remembered for the work he did to raise awareness globally of the plight facing not only rhinos, but also the many thousands of other species facing extinction. as a result of unsustainable human activity.
'One day, his demise will hopefully be seen as a seminal moment for conservationists world wide.'
Photograph: Mike Segar/Reuters.
