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England got off to a poor start in their second innings after Australia paceman Peter Siddle destroyed the top order. Siddle picked three quick wickets to have England end the day at 31-3 (20 overs) at stumps.
England have a 264-run lead with Joe Root (18 off 60) and Tim Bresnan (0 off 15) at the crease.
Earlier, Graeme Swann played a leading hand with bat and ball as Australia were shot out for 128 in 53.3 overs in the first innings on Day 2 of the second Ashes Test at Lord’s on Friday.
The off-spinner, who earlier in the day swelled England’s total with a quick 28 not out batting at No 11, had figures of five wickets for 44 runs.
Tim Bresnan, who took two wickets for nine runs to spark an initial collapse that saw Australia's 42 without loss transformed into 53 for three early in the second session, finished with 2 for 28.
Earlier in the day, Ryan Harris took five wickets (5-72), but late resistance helped England amass 361 in their first innings.
The 34-year-old, who replaced Mitchell Starc in the attack ahead of the Test, justified the selector's faith in him.
It was Harris's third career five-wicket haul, after his six for 47 against England at WACA in December 2010 and five for 62 against Sri Lanka at Galle in August 2011.
Harris struck with the day's first ball, having Tim Bresnan (7) caught behind. In his third over, the fifth of the day, he also had James Anderson (12) out in a similar fashion to complete his fiver.
The wicket of Jonathan Trott, his third in the innings, also happened to be his 50th Test wicket.
Former England captain Mike Brearley rang the bell before the start of play on Day 2.
Considered one of the best captains ever, Brearley had outstanding record in Ashes.
In 1981, he replaced Ian Botham as captain in the third Test at Headingley with England 0-1 down, and proceeded to win the series 3-1.
Stuart Broad (33) and Graeme Swann (28 not out) frustrated Australia, their last wicket stand worth 49 runs, and took the home team past the 350-run mark.
In the course of their partnership the duo played some fearless cricket, hitting all over the park.
James Pattinson eventually had Broad caught behind to pick his first wicket of the innings and end Australian misery.
Australia’s opener Shane Watson and Chris Rogers started the chase positively. The former, who has scored just two hundreds in 42 previous Tests, looked in good touch while scoring 30.
But with just two balls to go for lunch, he played across the line of a Tim Bresnan delivery and was given out lbw by Sri Lankan umpire Kumar
Dharmasena.
Even though wasted reviews had cost Australia dear during Ashes-holders England's 14-run first Test win at Trent Bridge last week, Watson sought a referral.
Third umpire Tony Hill, however, confirmed the decision, leaving Chris Rogers 12 unbeaten on his Middlesex home ground at the break.
Soon after lunch Rogers was lbw in bizarre fashion. The left-hander missed a pull at a Swann full toss after the ball appeared to slip out of the bowler's hand, and was 'boxed'.
South African umpire Marais Erasmus gave Rogers out lbw for 15 on the left-hander's Middlesex home ground.
That started the slide and Australia were in deep trouble after their captain Michael Clarke was dismissed, undone by an excellent full length delivery from Stuart Broad that had him lbw as he played down the wrong line.
The last thing Australia needed now was a run-out. But that is what they got when a terrible mix-up saw non-striker Ashton Agar charge down the pitch and back again as he failed to beat Matt Prior's throw to James Anderson.
With a 233-run lead England openers Alastair Cook and Joe Root came out to bat for their second innings.
Both batsmen played Ryan Harris and Shane Watson cautiously. But it was first-change bowler Peter Siddle who put England in the doldrums.
In the 11th over, he first had Cook bowled for 8 and soon enough had Jonathan Trott dismissed for a duck, chopping the ball on to his stumps.
Kevin Pietersen came to the crease, smashed a boundary and was later caught by Chris Rogers at point off Siddle.
Root and Bresnan then ensured no more damage was done and England were 31-3 at stumps.